Personally, I believe it's mostly the combination of
- less pixel fillrate
- less memory
- less mem bandwidth.
All of these Gigabyte tests were done at 1920x1200 with 8x(!)AA, for example I'm rather sure the bad result in Unigine was not just because of the halved geometry performance, but also because 768MB simply aren't enough with these settings. Tesselation is a rather memory-hungry feature itself, combine this with a high resolution and high level of AA, and exceeding 768MB should be easy.
I expect the 1GB version to be quite a bit faster at high resolutions with 8xAA.